K. Kashefi and D. R. Lovley, Extending the upper temperature limit for life, Science, vol.301, 2003.

K. Takai, K. Nakamura, T. Toki, U. Tsunogai, M. Miyazaki et al., Cell proliferation at 122 ? C and isotopically heavy CH 4 production by a hyperthermophilic methanogen under high-pressure cultivation, Proc. Natl. Acad. Sci, vol.105, pp.10949-10954, 2008.

J. P. Amend and A. Teske, Expanding frontiers in deep subsurface microbiology, Palaeogeogr. Palaeoclimatol. Palaeoecol, vol.219, pp.131-155, 2005.

E. Miranda-tello, M. Fardeau, P. Thomas, F. Ramirez, L. Casalot et al., Petrotoga mexicana sp. nov., a novel thermophilic, anaerobic and xylanolytic bacterium isolated from an oil-producing well in the Gulf of Mexico, Int. J. Syst. Evol. Microbiol, vol.54, pp.169-174, 2004.

U. Szewzyk, R. Szewzyk, T. A. Stenström, and . Thermophilic, anaerobic bacteria isolated from a deep borehole in granite in Sweden, Proc. Natl. Acad. Sci, vol.91, pp.1810-1813, 1994.

Y. M. Bar-on, R. Phillips, and R. Milo, The biomass distribution on Earth, Proc. Natl. Acad. Sci, 2018.

C. Magnabosco, L. Lin, H. Dong, M. Bomberg, H. Ghiorse et al., The biomass and biodiversity of the continental subsurface, Nat. Geosci, vol.11, pp.707-717, 2018.

J. K. Fredrickson and D. L. Balkwill, Geomicrobial processes and biodiversity in the deep terrestrial subsurface, Geomicrobiol. J, vol.23, pp.345-356, 2006.

A. Oren, Bioenergetic aspects of halophilism. Microbiol, Mol. Biol. Rev, vol.63, pp.334-348, 1999.

L. R. Krumholz, J. P. Mckinley, G. A. Ulrich, and J. M. Suflita, Confined subsurface microbialcommunities in Cretaceous rock, Nature, vol.386, pp.64-66, 1997.

M. Magot, B. Ollivier, and B. K. Patel, Microbiology of petroleum reservoirs, Antonie Leeuwenhoek, vol.77, pp.103-116, 2000.

K. Pedersen, Exploration of deep intraterrestrial microbial life: Current perspectives, FEMS Microbiol. Lett, vol.185, pp.9-16, 2000.

S. D'hondt, B. B. Jørgensen, D. J. Miller, A. Batzke, R. Blake et al., Distributions of microbial activities in deep subseafloor sediments, Science, vol.306, pp.2216-2221, 2004.

E. F. Delong, Microbial life breathes deep, Science, vol.306, pp.2198-2200, 2004.

O. Basso, J. Lascourrèges, M. Jarry, and M. Magot, The effect of cleaning and disinfecting the sampling well on the microbial communities of deep subsurface water samples, Environ. Microbiol, vol.7, pp.13-21, 2005.

M. Fardeau, F. Goulhen, M. Bruschi, N. Khelifi, J. Cayol et al., Archaeoglobus fulgidus and Thermotoga elfii, thermophilic isolates from deep geothermal water of the Paris Basin, Geomicrobiol. J, vol.26, pp.119-130, 2009.
URL : https://hal.archives-ouvertes.fr/hal-00512519

T. C. Onstott, D. P. Moser, S. M. Pfiffner, J. K. Fredrickson, F. J. Brockman et al., Indigenous and contaminant microbes in ultradeep mines, Environ. Microbiol, vol.5, pp.1168-1191, 2003.

L. Purkamo, M. Bomberg, M. Nyyssönen, I. Kukkonen, L. Ahonen et al., Heterotrophic communities supplied by ancient organic carbon predominate in deep Fennoscandian bedrock fluids, Microb. Ecol, vol.69, pp.319-332, 2015.

T. Onstott, T. J. Phelps, F. Colwell, D. Ringelberg, D. White et al., Observations pertaining to the origin and ecology of microorganisms recovered from the deep subsurface of Taylorsville Basin, Virginia. Geomicrobiol. J, vol.15, pp.353-385, 1998.

G. Zhang, H. Dong, Z. Xu, D. Zhao, and C. Zhang, Microbial diversity in ultra-high-pressure rocks and fluids from the chinese continental scientific drilling project in China, Appl. Environ. Microbiol, vol.71, pp.3213-3227, 2005.

F. S. Colwell, T. C. Onstott, M. E. Delwiche, D. Chandler, J. K. Fredrickson et al., Microorganisms from deep, high temperature sandstones: Constraints on microbial colonization, FEMS Microbiol. Rev, vol.20, pp.425-435, 1997.

P. Landrein, G. Vigneron, V. Delay, P. Lebon, M. Pagel et al., hydrodynamisme et thermicité dans le système sédimentaire multicouche recoupé par les forages Andra de Montiers-sur-Saulx (Meuse). Bull. de la Société Géologique de Fr, vol.184, pp.519-543, 2013.

M. Brunet and V. Le-pichon, Subsidence of the Paris Basin, J. Geophys. Res, vol.87, pp.8547-8560, 1982.

S. Poulain, C. Sergeant, M. Simonoff, C. Le-marrec, and S. Altmann, Microbial investigations in Opalinus Clay, an argillaceous formation under evaluation as a potential host rock for a radioactive waste repository, Geomicrobiol. J, vol.25, pp.240-249, 2008.

. Nf-t, Water Quality-Determination of pH, pp.90-98, 2001.

, ISO 11732 2005. Water Quality-Measurement of Ammonium-Method by Flow Analysis, 2005.

, Water Quality-Determination of Alkalinity-Part 1: Determination of Total and Composite Alkalinity, pp.9963-9964, 1994.

, Water Quality-Measurement of Dissolved Anions by Ionic Chromatography in Liquid Phase-Part 1: Measurement of Bromide, Chloride, Fluoride, Nitrate, Nitrite, Phosphate and Sulphate, pp.10304-10305, 2010.

, Atomic Absorption Spectrometry-Use Introduction, 1975.

Y. Copard, C. Di-giovanni, T. Martaud, P. Albéric, and J. Olivier, Using Rock-Eval 6 pyrolysis for tracking fossil organic carbon in modern environments: Implications for the roles of erosion and weathering, Earth Surf. Process. Landf, vol.31, pp.135-153, 2006.
URL : https://hal.archives-ouvertes.fr/hal-00022590

, EN ISO 11885. Water Quality-Determination of Selected Elements by Inductively Coupled Plasma Optical Emission Spectrometry

, AFNOR, 2009.

P. Blanc, A. Lassin, P. Piantone, M. Azaroual, N. Jacquemet et al., A geochemical database focused on low temperature water/rock interactions and waste materials, Appl. Geochem, vol.27, pp.2107-2116, 2012.
URL : https://hal.archives-ouvertes.fr/hal-00846739

D. L. Parkhurst and C. A. Appelo, User's Guide to Phreeqc (Version 2)-A Computer Program for Speciation, Batch-Reaction, One-Dimensional Transport, and Inverse Geochemical Calculations, Water-Resources Investigations Report

T. L. Miller and M. J. Wolin, A serum bottle modification of the Hungate technique for cultivating obligate anaerobes, Appl. Microbiol, vol.27, pp.985-987, 1974.

J. Kallmeyer, T. G. Ferdelman, K. H. Jansen, and B. B. Jørgensen, A high-pressure thermal gradient block for investigating microbial activity in multiple deep-sea samples, J. Microbiol. Methods, vol.55, pp.165-172, 2003.

A. Lerner, Y. Shor, A. Vinokurov, Y. Okon, and E. Jurkevitch, Can denaturing gradient gel electrophoresis (DGGE) analysis of amplified 16s rDNA of soil bacterial populations be used in forensic investigations?, Soil Biol. Biochem, vol.38, pp.1188-1192, 2006.

D. J. Lane, 16S/23S rRNA sequencing, Nucleic Acid Techniques in Bacterial Systematics

E. Stackebrandt and M. Goodfellow, , pp.130-141, 1991.

H. Schäfer and G. Muyzer, Denaturing gradient gel electrophoresis in marine microbial ecology, vol.30, pp.425-468, 2001.

G. C. Baker, J. J. Smith, and D. A. Cowan, Review and re-analysis of domain-specific 16S primers, J. Microbiol. Methods, vol.55, pp.541-555, 2003.

E. Zumstein, R. Moletta, and J. Godon, Examination of two years of community dynamics in an anaerobic bioreactor using fluorescence polymerase chain reaction (PCR) single-strand conformation polymorphism analysis, Environ. Microbiol, vol.2, pp.1462-2920, 2000.

T. Huber, G. Faulkner, and P. Hugenholtz, Bellerophon: A program to detect chimeric sequences in multiple sequence alignments, Bioinformatics, vol.20, pp.2317-2319, 2004.

T. Hall, BioEdit: A user-friendly biological sequence alignment editor and analysis program for Windows 95/98/NT, Nucleic Acids Symp. Ser, vol.41, pp.95-98, 1999.

T. H. Jukes and C. R. Cantor, Evolution of protein molecules, In Mammalian Protein Metabolism

H. N. Munro and . Ed, , pp.21-132, 1969.

N. Saitou and M. Nei, The neighbor-joining method: A new method for reconstructing phylogenetic trees, Mol. Biol. Evol, vol.4, pp.406-425, 1987.

J. Felsenstein, Confidence limits on phylogenies: An approach using the bootstrap, Evolution, vol.39, pp.783-791, 1985.

S. Kumar, G. Stecher, M. Li, C. Knyaz, K. Tamura et al., Molecular Evolutionary Genetics Analysis across computing platforms, Mol. Biol. Evol, vol.35, pp.1547-1549, 2018.

A. J. Katz and A. H. Thompson, Quantitative prediction of permeability in porous rock, Phys. Rev. Ser. B, vol.34, pp.8179-8191, 1986.

R. Rebeix, C. Le-gal-la-salle, P. Verdoux, S. Gianesinni, J. Lancelot et al., Tracing the origin of water and solute transfers in deep groundwater from Oxfordian, Dogger and Trias formations in the east of the Paris Basin-France, Phys. Chem. Earth, vol.36, pp.1496-1510, 2011.
URL : https://hal.archives-ouvertes.fr/hal-00593583

H. A. Barton, N. M. Taylor, B. R. Lubbers, and A. C. Pemberton, DNA extraction from low-biomass carbonate rock: An improved method with reduced contamination and the low-biomass contaminant database, J. Microbiol. Methods, vol.66, pp.21-31, 2006.

E. Gérard, D. Moreira, P. Philippot, M. J. Van-kranendonk, and P. López-garcía, Modern subsurface bacteria in pristine 2.7 Ga-old fossil xtromatolite drillcore samples from the fortescue group, PLoS ONE, 2009.

C. Moissl, S. Osman, M. T. La-duc, A. Dekas, E. Brodie et al., Molecular bacterial community analysis of clean rooms where spacecraft are assembled, FEMS Microbiol. Ecol, vol.61, pp.509-521, 2007.

S. O. Hashim, O. Delgado, R. Hatti-kaul, F. J. Mulaa, and B. Mattiasson, Starch hydrolysing Bacillus halodurans isolates from a Kenyan soda lake, Biotechnol. Lett, vol.26, pp.823-828, 2004.

M. Ulfah, I. Helianti, B. Wahyuntari, and N. Nurhayati, Characterization of a new thermoalkalophilic xylanase-producing bacterial strain isolated from Cimanggu hot spring, west Java, Indonesia. Microbiol. Indones, vol.5, pp.139-143, 2012.

A. Reysenbach, M. Ehringer, and K. Hershberger, Microbial diversity at 83 ? C in Calcite Springs, Another environment where the Aquificales. Extremophiles, vol.4, pp.61-67, 2000.

D. P. Chandler, F. J. Brockman, and J. K. Fredrickson, Use of 16S rDNA clone libraries to study changes in a microbial community resulting from ex situ perturbation of a subsurface sediment, FEMS Microbiol. Rev, vol.20, pp.217-230, 1997.

G. M. Luna, A. Dell'anno, and R. Danovaro, DNA extraction procedure: A critical issue for bacterial diversity assessment in marine sediments, Environ. Microbiol, vol.8, pp.308-320, 2006.

F. Martin-laurent, L. Philippot, S. Hallet, R. Chaussod, J. C. Germon et al., DNA extraction from soils: Old bias for new microbial diversity analysis methods, Appl. Environ. Microbiol, vol.67, pp.2354-2359, 2001.

T. Okubo, T. Tsukui, H. Maita, S. Okamoto, K. Oshima et al., Complete genome sequence of Bradyrhizobium sp. S23321: Insights into symbiosis evolution in soil oligotrophs, Microbes Environ, vol.27, pp.306-315, 2012.

J. R. Marchesi, A. J. Weightman, B. A. Cragg, R. J. Parkes, and J. C. Fry, Methanogen and bacterial diversity and distribution in deep gas hydrate sediments from the Cascadia Margin as revealed by 16S rRNA molecular analysis, FEMS Microbiol. Ecol, vol.34, pp.221-228, 2001.

M. Mergeay, S. Monchy, T. Vallaeys, V. Auquier, A. Benotmane et al., Ralstonia metallidurans, a bacterium specifically adapted to toxic metals: Towards a catalogue of metal-responsive genes, FEMS Microbiol. Rev, vol.27, pp.385-410, 2003.

A. Tiemeyer, H. Link, and D. Weuster-botz, Kinetic studies on autohydrogenotrophic growth of Ralstonia eutropha with nitrate as terminal electron acceptor, Appl. Microbiol. Biotechnol, vol.76, pp.75-81, 2007.

B. E. Grey and T. R. Steck, The viable but nonculturable state of Ralstonia solanacearum may be involved in long-term survival and plant infection, Appl. Environ. Microbiol, vol.67, pp.3866-3872, 2001.

W. M. Chen, N. Cho, S. Yang, A. Arun, C. Young et al., Aquabacterium limnoticum sp. nov., isolated from a freshwater spring, Int. J. Syst. Evol. Microbiol, vol.62, pp.698-704, 2012.

S. Kalmbach, W. Manz, J. Wecke, and U. Szewzyk, Aquabacterium gen. nov., with description of Aquabacterium citratiphilum sp. nov., Aquabacterium parvum sp. nov. and Aquabacterium commune sp. nov., three in situ dominant bacterial species from the Berlin drinking water system, Int. J. Syst. Bacteriol, vol.49, pp.769-777, 1999.

C. Simon, A. Wiezer, A. W. Strittmatter, and R. Daniel, Phylogenetic diversity and metabolic potential revealed in a glacier ice metagenome, Appl. Environ. Microbiol, vol.75, pp.7519-7526, 2009.

C. M. Santelli, B. N. Orcutt, E. Banning, W. Bach, C. L. Moyer et al., Abundance and diversity of microbial life in ocean crust, Nature, vol.453, pp.653-656, 2008.

X. Gong, Z. Liu, P. Guo, C. Chi, J. Chen et al., Bacteria in crude oil survived autoclaving and stimulated differentially by exogenous bacteria, PLoS ONE, vol.7, 2012.

B. C. Christner, G. Royston-bishop, C. M. Foreman, B. R. Arnold, M. Tranter et al., Limnological conditions in subglacial lake Vostok, Antarctica. Limnol. Oceanogr, vol.51, pp.2485-2501, 2006.

J. A. Colquhoun, S. C. Heald, L. Li, J. Tamaoka, C. Kato et al., Taxonomy and biotransformation activities of some deep-sea actinomycetes, Extremophiles, vol.2, pp.269-277, 1998.

V. V. Kevbrin, K. Zengler, A. M. Lysenko, and J. Wiegel, Anoxybacillus kamchatkensis sp. nov., a novel thermophilic facultative aerobic bacterium with a broad pH optimum from the Geyser valley, Kamchatka. Extremophiles, vol.9, pp.391-398, 2005.

E. Willerslev, A. Cooper, . Ancient, and . Dna, Proc. R. Soc. B, vol.272, pp.3-16, 2005.

X. Lin, D. Kennedy, J. Fredrickson, B. Bjornstad, and A. Konopka, Vertical stratification of subsurface microbial community composition across geological formations at the Hanford Site, Environ. Microbiol, vol.14, pp.414-425, 2012.

E. A. Eloe, C. N. Shulse, D. W. Fadrosh, S. J. Williamson, E. E. Allen et al., Compositional differences in particle-associated and free-living microbial assemblages from an extreme deep-ocean environment, Environ. Microbiol. Rep, vol.3, pp.449-458, 2011.

D. Elhottová, V. Kri?t?fek, J. Frouz, A. Nováková, and A. Chro?áková, Screening for microbial markers in Miocene sediment exposed during open-cast brown coal mining, Antonie Leeuwenhoek, vol.89, pp.459-463, 2006.

M. S. Wilson, J. B. Herrick, C. O. Jeon, D. E. Hinman, and E. L. Madsen, Horizontal transfer of phnAc dioxygenase genes within one of two phenotypically and genotypically distinctive naphthalene-degrading guilds from adjacent soil environments, Appl. Environ. Microbiol, vol.69, pp.2172-2181, 2003.

P. Verma, P. K. Pandey, A. K. Gupta, H. J. Kim, K. S. Baik et al., Shewanella indica sp. nov., isolated from sediment of the Arabian Sea, Int. J. Syst. Evol. Microbiol, vol.61, pp.2058-2064, 2011.

J. Z. Kaye, M. C. Márquez, A. Ventosa, and J. A. Baross, Halomonas neptunia sp. nov., Halomonas sulfidaeris sp. nov., Halomonas axialensis sp. nov. and Halomonas hydrothermalis sp. nov.: Halophilic bacteria isolated from deep-sea hydrothermal-vent environments, Int. J. Syst. Evol. Microbiol, vol.54, pp.499-511, 2004.

A. Lauer, K. B. Sørensen, and A. Teske, Phylogenetic Characterization of Marine Benthic Archaea in Organic-Poor Sediments of the Eastern Equatorial Pacific Ocean (ODP Site 1225). Microorganisms, vol.4, 2016.

Y. A. Frank, V. V. Kadnikov, S. N. Gavrilov, D. Banks, A. L. Gerasimchuk et al., Stable and Variable Parts of Microbial Community in Siberian Deep Subsurface Thermal Aquifer System Revealed in a Long-Term Monitoring Study, Front. Microbiol, 2016.

M. Bomberg, M. Nyyssönen, A. Nousiainen, J. Hultman, L. Paulin et al., Evaluation of Molecular Techniques in Characterization of Deep Terrestrial Biosphere, Open J. Ecol, vol.4, pp.468-487, 2014.

G. E. Jenneman, M. J. Mcinerney, and R. M. Knapp, Microbial penetration through nutrient-saturated berea sandstone, Appl. Environ. Microbiol, vol.50, pp.383-391, 1985.

K. Alain and J. Querellou, Cultivating the uncultured: Limits, advances and future challenges, Extremophiles, vol.13, pp.583-594, 2009.
URL : https://hal.archives-ouvertes.fr/hal-00561362

B. Ollivier and D. Alazard, The oil reservoir ecosystem, Handbook of Hydrocarbon Microbiology: Microbial Interactions with Hydrocarbons, Oils, Fats and Related Hydrophobic Substrates and Products, pp.2262-2268, 2010.

T. J. Mcgenity, R. T. Gemmell, W. D. Grant, and H. Stan-lotter, Origins of halophilic microorganisms in ancient salt deposits, Environ. Microbiol, vol.2, pp.243-250, 2000.

T. Blaise, J. Barbarand, M. Kars, F. Ploquin, C. Aubourg et al., Reconstruction of low burial (<100 ? C) in sedimentary basins: A comparison of geothermometer sensibility in the intracontinental Paris Basin. Marine Petroleum Geol, vol.53, pp.71-87, 2014.

A. Wilhelm, S. R. Larter, I. Head, P. Farrimond, R. Di-primio et al., Biodegradation of oil in uplifted basins prevented by deep-burial sterilization, Nature, vol.411, pp.1034-1037, 2001.

K. Yamane, H. Maki, T. Nakayama, T. Nakajima, N. Nomura et al., Diversity and similarity of microbial communities in petroleum crude oils produced in Asia, Biosc. Biotechnol. Biochem, vol.72, pp.2831-2839, 2008.

S. A. Fish, T. J. Shepherd, T. J. Mcgenity, and W. D. Grant, Recovery of 16S ribosomal RNA gene fragments from ancient halite, Nature, vol.417, pp.432-436, 2002.

G. Gales, N. Tsesmetzis, I. Neria, D. Alazard, S. Coulon et al., Preservation of ancestral Cretaceous microflora recovered from a hypersaline oil reservoir, Sci. Rep, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01443597

F. Inagaki, H. Okada, A. I. Tsapin, and K. H. Nealson, Microbial survival: The Paleome: A Sedimentary Genetic Record of Past Microbial Communities, Astrobiology, vol.5, pp.141-153, 2005.

B. Marty, S. Dewonck, and C. France-lanord, Geochemical evidence for efficient aquifer isolation over geological timeframes, Nature, vol.425, pp.55-58, 2003.

H. Celle-jeanton, F. Huneau, Y. Travi, and W. M. Edmunds, Twenty years of groundwater evolution in the Triassic sandstone aquifer of Lorraine: Impacts on baseline water quality, Appl. Geochem, vol.24, pp.1198-1213, 2009.
URL : https://hal.archives-ouvertes.fr/hal-00453278

J. C. Fontes and J. M. Matray, Geochemistry and origin of formation brines from the Paris Basin, France: 1. Brines associated with Triassic salts, Chem. Geol, vol.109, pp.149-175, 1993.